What is it about smiling that can turn an otherwise ordinary bad guy into a memorable villain? Smiles, we are told, reveal that someone is happy about something. A villain, though, uses a smile for a different purpose: not to reveal, but to conceal. A great movie villain often doesn't see himself as a villain, but nevertheless, he knows that the best way to fit in -- to deflect suspicion and give the appearance that he is anything but a villain -- is a simple smile.

Not too long ago, we took a look at 15 famous screen villainesses who hid some serious dark sides beneath sunny, smiley exteriors. Now, it's time for the men -- and we bet you can't guess who comes in at No. 1. (Hint: He's not human.)

The 20 Smilingest Male Villains in Movies

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Photos courtesy of Everett Collection, Miramax, Fox.